The Wardens of Evolution, simply called as Wardens, is a forced role bestowed by the Great Tree of Origin. Particularly, these individuals were the last survivors of their kind who attained immortality. Cursed to see their civilization at its doom, they also had no choice but to see the other civilizations end.

Due to the Great Tree's Selection and Rejection process, called Lacrimosa, to watch the world after it wipes out all but one races from the world. The most important requirement to become one of these is to have a pure and bright soul. An insignia will be placed on the chosen's body as a symbol that they were chosen. Each of the Wardens once fought to stop the Lacrimosa but ended up accepting it in the end, as fighting it would be futile.

Usually before a Lacrimosa is descended to a race, the previous chosen Warden will start searching for the next warden. The exceptions are: Hydra and Adol. Hydra, being the first warden, was selected by the Tree directly. Adol was connected with the last warden, Dana, but Dana did not search for him like Ura did to her.

During the true ending, all of the Wardens were freed from their insignia and decided to help Dana watch the new world, while Adol will keep going on various adventures.

All wardens has the Insignia of Warden marked somewhere on their bodies. Hydra on the back of his right hand. Minos on his lower left belly. Nestor on her right side of her neck. Dana and Adol both have their marks on their left shoulders. Ura is the only one whose Insignia was not shown; however, Ricotta's mentioning "All of them has the same mark as Adol and Dana!" suggests that Ura has it somewhere visible.

Members[]

Hydra[]

A tall man resembling a marine creature. He leads his marine species to prosperity but got chosen at the start of their era's Lacrimosa. He is the one who built the Seren Garden and the Tree of Psyches situated at the Valley of Kings. He is the one who knew the tree's ability to stop the Great Tree's Lacrimosa but the psyche of their species was not enough to stop it.

Minos[]

A tall muscled beastlike being whose appearance is indicating his rugged temperament. He lead his species during the Ice era as their king. As the Lacrimosa hit their civilization, he discovered the Tree of Psyches and tried to use it to stop the Lacrimosa. But in the end the accumulated psyche was not enough to grow the tree into its maturity.

Nestor[]

A woman with insect like features. She also lead her species as Queen until she was chosen as Warden and her world destroyed by the Lacrimosa. She also discovered the Tree of Psyches.

Ura[]

A woman who mimicked the appearance of the Eternian Princess, Sarai. Her people descended from the skies and used their power of mimicry to claim dominion over their world. However their civilization was also destroyed, leaving her as the last member of her race.

Dana Iclucia[]

The Maiden of the Great Tree during the Eternian Era. Her species evolved from Saurians, later known as Primordials, making them tall and powerful. She became a warden after her fight with Le-Kyanos, the mysterious Primordial she encountered in the fog in front of the Great Tree of Origins. She also tried to stop the Lacrimosa, but the impending calamity and weather abnormalities after the meteor strike almost made her surrender. However she sealed herself at the base of the Great Tree and became known as a natural distortion to the world's cycle along with Adol. Later, she ascended and become the Goddess of Evolution, with the other wardens as her assistants.

Adol Christin[]

The main protagonist of the game. He became involved with the Lacrimosa because of his dreams about Dana's past. He was chosen after he defeated Le-Erythros inside the fog.